Invertebrate Zoology

Associate Curator Emeritus Chen W. Young Chen Young
412.688.8670 (fax)

Click here for full bio 

Academic Training 

PhD, University of Kansas, 1980
BS, National Chung-Hsing University, Taiwan, 1971


Responsibilities include conducting original and focused field work involving collection and natural history research for the purpose of enhancing the scientific and educational merit of the museum's entomological programs. Provide assistance to Division of Public Programs for the maintenance and development of programs and exhibitions.

General Research Interests 

  • Systematics of crane flies, the Tipuloidea (Diptera: Nematocera)
  • Field-oriented studies on natural history of Tipulidae, including larval morphology, ontogeny, ecology and habitat relations, faunistic studies
  • Systematic and evolutionary studies,& including historical biogeography
  • Systematics and natural history of Neotropical Lymantriidae (Lepidoptera: Noctuoidea)

Web Sites 

Research Emphases 

  • Biodiversity of crane flies in Taiwan
  • Revisionary treatment of the crane fly genus Dapanoptera of Australian-Oceanian regions
  • Revision of crane fly genus Megistomastix in Hispaniola
  • Systematics and biogeography of Tipulidae (crane flies) in the Caribbean region
  • Flightless crane flies in the Ecuadoran Andes
  • Immature stages of Microtipula, Eumicrotipula, Zelandotipula, Bellardina in Tipulinae
  • Survey of the Tipulidae of Powdermill Nature Reserve, the environmental research center of Carnegie Museum of Natural History, and the state of Pennsylvania (with J.K. Gelhaus)
  • Fauna of Tipulidae in Kyrgyzstan and in China (with D. Yang)
  • Revision of Neotropical gypsy moths (Lymantriidae) in the genera Sarsina and Staetherinia (with J.E. Rawlins)
  • Synoptic work on the macrolepidopterous larvae of Taiwan and eastern North America (with J.E. Rawlins)

Selected Publications 

  • 2010 Larva, pupa and biology of the crane fly Indotipula demarcata (Brunetti) from Taiwan (Insecta: Diptera: Tipulidae. Annals of Carnegie Museum, 78(4):319-324 (with C. Chu). Click here for e-print (PDF, 1.2 Mb).
  • 2009 New species and immature of crane flies of subgenus Formotipula Matsumura from Taiwan (Diptera: Tipulidae: Tipula). Zoosymposia, 3:309-321. Click here for e-print (PDF, 2.2 Mb).
  • 2008 Global diversity of crane flies (Insecta: Diptera: Tipulidae or Tipulidae sensu lato) in fresh water. Hydrobiologia, 595:457-467. (with H deJong, P. Oosterbroek, J. Gelhaus, H. Resch). Click here for e-print (PDF, 322 Kb).
  • 2007 Notes on crane fly species of Prionota Van Der Wulp from China (Diptera: Tipulidae). Annals of Carnegie Museum, 76(3):165-170. Click here for e-print (PDF, 1.7 Mb).
  • 2007 First observation of parasitic relations between big-headed flies, Nephrocerus Zetterstedt (Diptera: Pipunculidae) and crane flies, Tipula (Diptera: Tipulidae: Tipulinae), with larval and pupal descriptions for the genus Nephrocerus. Proceedings of the Entomological Society of Washington, 109(1):52-65. (with D. Koenig). Click here for e-print (PDF, 1.2 Mb).
  • 2006 Diptera: Tipulidae. In Li Z. Z. et al (eds.) Insects from Fanjingshan landscape. Guizhou Science and Technology Publishing House, Guiyang, Guizhou, China, 445-449.
  • 2004 Family Tipulidae. in C. Yule, et al (eds.) The Freshwater Invertebrates of Malaysia and Singapore. Academy of Sciences Malaysia, Kula Lumpur, Malaysia, 775-785. Click here to read more about this publication. 
  • 2003 Biology and immature stages of the crane fly Ptilogyna (Plusiomyia)herroni (Alexander) form New Caledonia, with discussion of its phylogenetic placement (Diptera: Tipulidae). Proceedings of the Entomological Society of Washington, 105(2):452-459 (with Hynes). Click here for e-print (PDF, 739 Kb).
  • 2002 Notes on female brachyptery in Nephrotoma basiflava Yang and Yang (Diptera: Tipulidae). Journal of Kansas Entomological Society 75(2):110-115 (with Yang). Click here to read more about this discovery. 
  • 2001 Leptotarsus (Tanypremna) in the Lesser Antilles: Description of a new species from Guadeloupe and biogeographical notes (Diptera: Tipulidae). Annals of Carnegie Museum, 70(4):239-245. Click here for e-print (PDF, 549 Kb).
  • 2000 Notes on the mature larva and pupa of Tipula (Lunatipula) georgiana Alexander (Diptera: Tipulidae). Transactions of the American Entomological Society, 126:417-421 (with D.P. Koenig). Click here for e-print (PDF, 310 Kb).
  • 2000 First occurrence of Tipula (Tipula) oleracea Linnaeus (Diptera: Tipulidae) in the New World with biological notes. Journal of the Kansas Entomological Society, 72(2):226-232 (with G. Onore & K. Proao). Click here for e-print (PDF, 511 Kb).
  • 1999 New species and immature instars of crane flies in the subgenus Tipulodina Enderlein from Sulawesi (Insecta: Diptera: Tipulidae: Tipula). Annals of Carnegie Museum, 68(2):81-90. Click here to read more about this discovery. 
  • 1995 Pupae of the crane fly genus Leptotarsus (Diptera: Tipulidae) in the New World, with discussion of the monophyly of the genus. Annals of Carnegie Museum, 64(2):135-145 (with J. Gelhaus). Click here for e-print (PDF, 833 Kb).
  • 1994 A new species of Antocha (subgenus Orimargula) from Sulawesi (Diptera: Tipulidae) and its mate-clasping behavior. Annals of Carnegie Museum, 63(4):319-325. Click here for e-print (PDF, 488 Kb).
  • 1992 Observations on the bizarre jelly mass habit of larval Geranomyia (Diptera: Tipulidae: Limoniinae). Journal of the New York Entomological Society, 100(4):634-637 (with D. Grimaldi).
  • 1992 Leptotarsus (Longurio) byersi, a new flightless crane fly from Ecuador (Diptera: Tipulidae). Acta Zoologica cracoviensa, 35(1):97-105 (with J. Gelhaus).Click here for e-print (PDF, 597 Kb).
  • 1991 The immature instars and biology of the crane fly genus Brachypremna Osten Sacken (Diptera: Tipulidae). Proceedings of the Entomological Society of Washington, 93(3):613-621 (with J. Gelhaus). Click here for e-print (PDF, 914 Kb).
  • 1990 A new Limonia (subgenus Laosa) species from Sulawesi (Dipt., Tipulidae) and its resting behaviour. Entomologist's Monthly Magazine, 126(1990):239-244. Click here for e-print (PDF, 491 Kb).
  • 1987 A review of the crane flies in the subgenus Tipula (Papuatipula) (Diptera: Tipulidae), with descriptions of five new species. Annals of Carnegie Museum, 56(7):161-171. Click here for e-print (PDF, 939 Kb).
  • 1987 A revision of the crane fly genus Dicranoptycha in North America. The University of Kansas Science Bulletin, 53(5):215-274