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Laonastes aenigmamus (recent) found in Khammouan Province, Laos, standing over the fossil relative Diatomys shantungensis (20 mya) from Shandong, China.
Illustration: Mark A. Klingler/CMNH

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Diatomys shantungensis fossil diagram
Illustration: Mark A. Klingler/CMNH

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Diatomys shantungensis fossil diagram
Photo: Mark A. Klingler/CMNH

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Scientists Mary Dawson and Laurent Marivaux with Laonastes skull specimens.
Photo: L. Marivaux/CNRS

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Laonastes skull and jaw, lateral view
Photo: L. Marivaux/CNRS

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Laonastes lower jaw specimen
Photo: L. Marivaux/CNRS

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Laonastes molar specimen depicting the four roots.
Photo: L. Marivaux/CNRS

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Laonastes skull
Photo: L. Marivaux/CNRS

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Laonastes skull maxilla specimen
Photo: L. Marivaux/CNRS

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Locality map for Diatomys shantungensis and Laonastes aenigmamus.
Map: Mark A. Klingler/CMNH

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Simplified tree: the phylogenetic relationships within the Hystricognathi.
Graphic: Mark A. Klingler/CMNH

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