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BeardCurator & Mary R. Dawson Chair of Vertebrate Paleontology

Ph.D., 1989, The Johns Hopkins University, School of Medicine

Phone: (412) 622-5782
Fax: (412) 622-5521; (412) 622-8837
Email: beardc@carnegiemnh.org
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Complete Bibliography
  • Kay, R. F., D. T. Rasmussen and K. C. Beard. 1984. Cementum annulus counts provide a means for age determination in Macaca mulatta (Primates, Anthropoidea). Folia Primatologica 42:85-95.

  • Beard, K. C., M. F. Teaford and A. Walker. 1986. New wrist bones of Proconsul africanus and P. nyanzae from Rusinga Island, Kenya. Folia Primatologica 47:97-118.

  • Beard, K. C. 1987. Jemezius, a new omomyid primate from the early Eocene of northwestern New Mexico. Journal of Human Evolution 16:457-468.

  • Beard, K. C., M. Dagosto, D. L. Gebo and M. Godinot. 1988. Interrelationships among primate higher taxa. Nature 331:712-714.

  • Beard, K. C. 1988. New notharctine primate fossils from the early Eocene of New Mexico and southern Wyoming and the phylogeny of Notharctinae. American Journal of Physical Anthropology 75:439-469.

  • Beard, K. C., and M. Godinot. 1988. Carpal anatomy of Smilodectes gracilis (Adapiformes, Notharctinae) and its significance for lemuriform phylogeny. Journal of Human Evolution 17:71-92.

  • Gebo, D. L., K. C. Beard, M. F. Teaford, A. Walker, S. G. Larson, W. L. Jungers and J. G. Fleagle. 1988. A hominoid proximal humerus from the early Miocene of Rusinga Island, Kenya. Journal of Human Evolution 17:393-401.

  • Beard, K. C. 1988. The phylogenetic significance of strepsirhinism in Paleogene primates. International Journal of Primatology 9:83-96.

  • Teaford, M. F., K. C. Beard, R. E. Leakey and A. Walker. 1988. New hominoid facial skeleton from the early Miocene of Rusinga Island, Kenya, and its bearing on the relationship between Proconsul nyanzae and Proconsul africanus. Journal of Human Evolution 17:461-477.

  • Beard, K. C., and P. Houde. 1989. An unusual assemblage of diminutive plesiadapiforms (Mammalia, ?Primates) from the early Eocene of the Clark's Fork Basin, Wyoming. Journal of Vertebrate Paleontology 9:388-399.

  • Bown, T. M., and K. C. Beard. 1990. Systematic lateral variation in the distribution of fossil mammals in alluvial paleosols, lower Eocene Willwood Formation, Wyoming. Pp. 135-151 in Dawn of the Age of Mammals in the Northern Part of the Rocky Mountain Interior, North America (T. M. Bown and K. D. Rose, eds.). Geological Society of America Special Paper, No. 243.

  • Beard, K. C. 1990. Gliding behaviour and palaeoecology of the alleged primate family Paromomyidae (Mammalia, Dermoptera). Nature 345:340-341.

  • Krishtalka, L., R. K. Stucky and K. C. Beard. 1990. The earliest fossil evidence for sexual dimorphism in primates. Proceedings of the National Academy of Sciences, USA 87:5223-5226.

  • Beard, K. C. 1990. Do we need the newly proposed order Proprimates? Journal of Human Evolution 19:817-820.

  • Beard, K. C., L. Krishtalka and R. K. Stucky. 1991. First skulls of the early Eocene primate Shoshonius cooperi and the anthropoid-tarsier dichotomy. Nature 349:64-67.

  • Beard, K. C., and A. R. Tabrum. 1991. The first early Eocene mammal from eastern North America: an omomyid primate from the Bashi Formation, Lauderdale County, Mississippi. Mississippi Geology 11 (2):1-6.

  • Beard, K. C., and B. Wang. 1991. Phylogenetic and biogeographic significance of the tarsiiform primate Asiomomys changbaicus from the Eocene of Jilin Province, People's Republic of China. American Journal of Physical Anthropology 85:159-166.

  • Beard, K. C. 1991. Vertical postures and climbing in the morphotype of Primatomorpha: implications for locomotor evolution in primate history. Pp. 79-87 in Origines de la Bipédie chez les Hominidés (Y. Coppens and B. Senut, eds.). Editions du CNRS (Cahiers de Paléoanthropologie), Paris.

  • Godinot, M., and K. C. Beard. 1991. Fossil primate hands: a review and an evolutionary inquiry emphasizing early forms. Human Evolution 6:307-354.

  • Beard, K. C., L. Krishtalka and R. K. Stucky. 1992. Revision of the Wind River faunas, early Eocene of central Wyoming. Part 12. New species of omomyid primates (Mammalia: Primates: Omomyidae) and omomyid taxonomic composition across the early-middle Eocene boundary. Annals of Carnegie Museum 61:39-62.

  • Beard, K. C., B. Sigé and L. Krishtalka. 1992. A primitive vespertilionoid bat from the early Eocene of central Wyoming. Comptes Rendus de l'Académie des Sciences, Paris (Série II) 314:735-741.

  • Beard, K. C. 1993. Phylogenetic systematics of the Primatomorpha, with special reference to Dermoptera. Pp. 129-150 in Mammal Phylogeny: Placentals (F. S. Szalay, M. J. Novacek and M. C. McKenna, eds.). Springer-Verlag, New York.

  • Preuschoft, H., M. Godinot, K. C. Beard, U. Nieschalk and F. K. Jouffroy. 1993. Biomechanical considerations to explain important morphological characters of primate hands. Pp. 245-256 in Hands of Primates (H. Preuschoft and D. J. Chivers, eds.). Springer-Verlag, Vienna.

  • Godinot, M., and K. C. Beard 1993. A survey of fossil primate hands. Pp. 335-378 in Hands of Primates (H. Preuschoft and D. J. Chivers, eds.). Springer-Verlag, Vienna.

  • Beard, K. C., M. F. Teaford and A. Walker. 1993. New hand bones of the early Miocene hominoid Proconsul and their implications for the evolution of the hominoid wrist. Pp. 387-403 in Hands of Primates (H. Preuschoft and D. J. Chivers, eds.). Springer-Verlag, Vienna.

  • Dawson, M. R., M. C. McKenna, K. C. Beard and J. H. Hutchison. 1993. An early Eocene plagiomenid mammal from Ellesmere and Axel Heiberg Islands, Arctic Canada. Kaupia 3:179-192.

  • Beard, K. C. 1993. Origin and evolution of gliding in early Cenozoic Dermoptera (Mammalia, Primatomorpha). Pp. 63-90 in Primates and Their Relatives in Phylogenetic Perspective (R. D. E. MacPhee, ed.). Plenum Press, New York.

  • Rose, K. D., K. C. Beard and P. Houde. 1993. Exceptional new dentitions of the diminutive plesiadapiforms Tinimomys and Niptomomys (Mammalia), with comments on the upper incisors of Plesiadapiformes. Annals of Carnegie Museum 62:351-361.

  • Beard, K. C., T. Qi, M. R. Dawson, B. Wang and C. Li. 1994. A diverse new primate fauna from middle Eocene fissure-fillings in southeastern China. Nature 368:604-609.

  • Beard, K. C., and R. D. E. MacPhee. 1994. Cranial anatomy of Shoshonius and the antiquity of Anthropoidea. Pp. 55-97 in Anthropoid Origins (J. G. Fleagle and R. F. Kay, eds.). Plenum Press, New York.

  • Beard, K. C., and J. Wang. 1995. The first Asian plesiadapoids (Mammalia: Primatomorpha). Annals of Carnegie Museum 64:1-33.

  • MacPhee, R. D. E., K. C. Beard and T. Qi. 1995. Significance of primate petrosal from middle Eocene fissure-fillings at Shanghuang, Jiangsu Province, People's Republic of China. Journal of Human Evolution 29:501-514.

  • Beard, K. C., Y. Tong, M. R. Dawson, J. Wang and X. Huang. 1996. Earliest complete dentition of an anthropoid primate from the late middle Eocene of Shanxi Province, China. Science 272:82-85.

  • Qi, T., and K. C. Beard. 1996. Nanotitan shanghuangensis, gen. et sp. nov.: the smallest known brontothere (Mammalia: Perissodactyla). Journal of Vertebrate Paleontology 16:578-581.

  • Qi, T., K. C. Beard, B. Wang, M. R. Dawson, J. Guo and C. Li. 1996. The Shanghuang mammalian fauna, middle Eocene of Jiangsu: History of discovery and significance. Vertebrata PalAsiatica 34:202-214.

  • Dawson, M. R., and K. C. Beard. 1996. New late Paleocene rodents (Mammalia) from Big Multi Quarry, Washakie Basin, Wyoming. Palaeovertebrata 25:301-321.

  • Beard, K. C. 1998. Biostratigraphy and paleobiogeography of Asian land mammals near the Paleocene-Eocene boundary: A phylogenetic approach. Strata: Actes du Laboratoire de Géologie Sédimentaire et Paléontologie de l'Université Paul-Sabatier (Toulouse, France) (Série 1) 9:25-28.

  • Beard, K. C. 1998. East of Eden: Asia as an important center of taxonomic origination in mammalian evolution. Pp. 5-39 in Dawn of the Age of Mammals in Asia (K. C. Beard and M. R. Dawson, eds.). Bulletin of Carnegie Museum of Natural History, No. 34.

  • Beard, K. C. 1998. A new genus of Tarsiidae (Mammalia: Primates) from the middle Eocene of Shanxi Province, China, with notes on the historical biogeography of tarsiers. Pp. 260-277 in Dawn of the Age of Mammals in Asia (K. C. Beard and M. R. Dawson, eds.). Bulletin of Carnegie Museum of Natural History, No. 34.

  • Qi, T., and K. C. Beard. 1998. Late Eocene sivaladapid primate from Guangxi Zhuang Autonomous Region, People's Republic of China. Journal of Human Evolution 35: 211–220.

  • Wilf, P., K. C. Beard, K. S. Davies-Vollum, and J. W. Norejko. 1998. Portrait of a late Paleocene (early Clarkforkian) terrestrial ecosystem: Big Multi Quarry and associated strata, Washakie Basin, southwestern Wyoming. Palaios 13: 514–532.

  • Dagosto, M., D. L. Gebo and K. C. Beard. 1999. Revision of the Wind River faunas, early Eocene of central Wyoming. Part 14. Postcranium of Shoshonius cooperi (Mammalia: Primates). Annals of Carnegie Museum 68:175-211.

  • Beard, K. C. and M. R. Dawson. 1999. Intercontinental dispersal of Holarctic land mammals near the Paleocene/Eocene boundary: paleogeographic, paleoclimatic and biostratigraphic implications. Bulletin de la Société Géologique de France 170:697-706.

  • Gebo, D. L., M. Dagosto, K. C. Beard, and J. Wang. 1999. A first metatarsal of Hoanghonius stehlini from the late middle Eocene of Shanxi Province, China. Journal of Human Evolution 37: 801-806.

  • Gebo, D. L., M. Dagosto, K.C. Beard, T. Qi, and J. Wang. 2000. The oldest known anthropoid postcranial fossils and the early evolution of higher primates. Nature 404: 276-278.

  • Gebo, D. L., M. Dagosto, K.C. Beard and T. Qi. 2000. The smallest primates. Journal of Human Evolution 38: 585-594.

  • Beard, K. C. 2000. A new species of Carpocristes (Mammalia: Primatomorpha) from the middle Tiffanian of the Bison Basin, Wyoming, with notes on carpolestid phylogeny. Annals of Carnegie Museum 69: 195-208.

  • Guo, J., Dawson, M. R., and Beard, K. C. 2000. Zhailimeryx, a new lophiomerycid artiodactyl (Mammalia) from the late middle Eocene of central China and the early evolution of ruminants. Journal of Mammalian Evolution 7: 239-258.

  • Beard, K. C., and Dawson, M. R. 2001. Early Wasatchian mammals from the Gulf Coastal Plain of Mississippi: Biostratigraphic and paleobiogeographic implications. Pp. 75-94 in Eocene Biodiversity: Unusual Occurrences and Rarely Sampled Habitats (G. F. Gunnell, ed.). Kluwer Academic/Plenum Publishers, New York.

  • Gebo, D. L., Dagosto, M., Beard, K. C., and Qi, T. 2001. Middle Eocene primate tarsals from China: Implications for haplorhine evolution. American Journal of Physical Anthropology 116: 83-107. E-print (709 Kb PDF)

  • Beard, K. C. 2002. East of Eden at the Paleocene/Eocene boundary. Science 295: 2028-2029.

  • Beard, K. C. 2002. Basal anthropoids. Pp. 133-149 in The Primate Fossil Record (W. C. Hartwig, ed.). Cambridge University Press, Cambridge, United Kingdom.

  • Beard, K. C., and Wang, J. 2004. The eosimiid primates (Anthropoidea) of the Heti Formation, Yuanqu Basin, Shanxi and Henan Provinces, People's Republic of China. Journal of Human Evolution 46: 401-432. E-print (1.9 Mb PDF)

  • Beard, K. C. 2004. The Hunt for the Dawn Monkey. The University of California Press, Berkeley, 348 pp. View publication info

  • Beard, K. C., Jaeger, J.-J., Chaimanee, Y., Rossie, J. B., Aung Naing Soe,
    Soe Thura Tun, Marivaux, L., and Marandat, B. 2005. Taxonomic status of
    purported primate frontal bones from the Eocene Pondaung Formation of
    Myanmar. Journal of Human Evolution 49: 468-481. E-print (1.2 Mb PDF)

  • Métais, G., Qi, T., Guo, J.-W., and Beard, K. C. 2005. A new bunoselenodont
    artiodactyl from the middle Eocene of China and the early record of
    selenodont artiodactyls in Asia. Journal of Vertebrate Paleontology 25:
    994-997.

  • Dawson, M. R., Marivaux, L., Li, C.-K., Beard, K. C., and Métais, G. 2006.
    Laonastes and the "Lazarus Effect" in Recent mammals. Science 311:
    1456-1458. E-print (493 Kb PDF)

  • Rossie, J. B., Ni, X.-J., and Beard, K. C. 2006. Cranial remains of an
    Eocene tarsier. Proceedings of the National Academy of Sciences 103:
    4381-4385. E-print (788 Kb PDF)

  • Beard, K. C. 2008. The oldest North American primate and mammalian biogeography during the Paleocene-Eocene Thermal Maximum. Proceedings of the National Academy of Sciences 105: 3815-3818. E-print (741 Kb PDF)

  • Marivaux, L., Beard, K. C., Chaimanee, Y., Jaeger, J.-J., Marandat, B., Aung Naing Soe, Soe Thura Tun, Htun Htun Aung, and Wanna Htoon. 2008. Anatomy of the bony pelvis of a relatively large-bodied strepsirrhine primate from the late middle Eocene Pondaung Formation (central Myanmar). Journal of Human Evolution 54: 391-404. E-print (826 Kb PDF)

  • Meng, J., Ni, X., Li, C., Beard, K. C., Gebo, D. L., Wang, Y., and Wang, H.
    2007. New material of Alagomyidae (Mammalia, Glires) from the late Paleocene Subeng locality, Inner Mongolia. American Museum Novitates 3597: 1-29.

  • Gebo, D. L., Dagosto, M., Beard, K. C., Ni, X., and Qi, T. 2007. Primate humeral remains from the middle Eocene of China. Bulletin of Carnegie Museum of Natural History 39: 77-82. E-print (130 Kb PDF)

  • Beard, K. C., Marivaux, L., Soe Thura Tun, Aung Naing Soe, Chaimanee, Y., Wanna Htoon, Marandat, B., Htun Htun Aung, and Jaeger, J.J. 2007. New sivaladapid primates from the Eocene Pondaung Formation of Myanmar and the anthropoid status of Amphipithecidae. Bulletin of Carnegie Museum of Natural History 39: 67-76. E-print (964 Kb PDF)

  • Dawson, M. R., and Beard, K. C. 2007. Rodents of the family Cylindrodontidae
    (Mammalia) from the earliest Eocene of the Tuscahoma Formation, Mississippi.
    Annals of Carnegie Museum 76: 135-144.
    E-print (2.1 Mb PDF)

  • Ni, X., Beard, K. C., Meng, J., Wang, Y., and Gebo, D. L. 2007. Discovery of the first early Cenozoic euprimate (Mammalia) from Inner Mongolia. American Museum Novitates 3571: 1-11. E-print (512 Kb PDF)

  • Meng, J., Wang, Y., Ni, X., Beard, K. C., Sun, C., Li, Q., Jin, X., and Bai, B. 2007. New stratigraphic data from the Erlian Basin: Implications for the division, correlation, and definition of Paleogene lithological units in Nei Mongol (Inner Mongolia). American Museum Novitates 3570: 1-31. E-print (3.3 Mb PDF)

  • Meng, J., Li, C., Ni, X., Wang, Y., and Beard, K. C. 2007. A new Eocene rodent from the lower Arshanto Formation in the Nuhetingboerhe (Camp Margetts) area, Inner Mongolia. American Museum Novitates 3569: 1-18. E-print (2.8 Mb PDF)
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